In the front, I originally put the all weather mats on top of the carpet mats. Later I removed and stored the front carpet mats. I don't think it hurt them to be underneath, I just didn't like the added height.
For the rears, I left the carpet mats underneath. After 10 months they still look pristine.
I removed my original mats and replaced them with the weathertechs. As for the tray, mine was delivered rolled up so I don't think it would do any harm to roll it up and put it behind the third row. I plan on doing this when I need to use the third row.
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